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alochthonous amphibians anatomía aragón ayuda azolla filiculoides basque country biotopes bullfrog cannibalism competence concolor morph conservation dispersal distribution emys orbicularis european pond turtle fishery food resources fumarole gallotia habitat conservation hatchling helminths histology historical presence iberian green frog iberian peninsula iberolacerta monticola intraspecific predation introduced species invasive species lacertid lacertidae lithobates catesbeianus loggerhead sea turtle longline mauremys leprosa mauremys rivulata melanism mesotriton alpestris montsant mountains muda natrix maura new records north-western iberia northwest spain nw spain ocular ophidia pelobates cultripes physalaemus biligonigerus podarcis hispanica pogona pomatias elegans population monitoring refuge relict population sea turtle sierra nevada snakes spain tadpole testis threatened populations tupinambis merianae turkey viperine snake viviparous lizard western sahara